The price of a peak time single ticket on the direct service from London Paddington to the airport increases from £22 to £24, and a business class ticket from £30 to £32.
The hike is part of an attempt to ease congestion on the line by encouraging off-peak travel.
Off-peak return tickets will soon be reduced from £36 to £25.
A single fare on the Heathrow Connect, which also departs from Paddington, costs £10.20, and travelling to the airport on the Underground from central London costs £5.10 at peak times.
Heathrow Express director Fraser Brown said that the changes would protect the ‘great service’ provided on the quickest route to Heathrow.
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